Darevsky, I.S. & Vedmederja, W.I. (1977) - A new species of Rock lizard, Lacerta saxicola EVERSMANN-Group from northeastern Turkey and adjoining regions of Adjaria. - Proceedings of the Zoological Institute of the USSR Academy of Sciences, Leningrad, 74: 50-54.  Schmidtler, J.F. & Heckes, U. & Bischoff, W. & Franzen, M. (2002) - Höhenabhängige Merkmalsvariation bei Felseidechsen des Darevskia clarkorum (DAREVSKY & VEDMEDERJA, 1977) / D. dryada (DAREVSKY & TUNIYEV, 1997)-Komplexes: Ein Fall von klimaparalleler Pholidosevariation? - Faunistische Abhandlungen Staatliches Museum für Tierkunde in Dresden, 23 (8): 141-156.  × External characters of 75 specimens of the nominal taxa Darevskia clarkorum (DAREVSKY & VEDMEDERJA, 1977) and Darevskia dryada (DAREVSKY & TUNIYEV, 1997) from between westernmost Georgia and the Turkish province of Giresun and from elevations between 250 m and 1750 m a. s. 1. were investigated by univariate and multivariate analysis. The pholidosis displays mainly two groups corresponding to samples from lower (colline) and higher (mon- tane) elevations. Samples from intermediate elevations show a somewhat intermediate pho- lidosis, but are apparently closer to higher elevation samples.
In general, these observations correspond with the results of DAREVSKY & TUNIYEV (1997). Based on this variation DAREVSKY & TUNIYEV (1997) separate two species, D. dryada from the colline zone, which they newly describe, and D. clarkorum, which they suppose to inhabit montane areas. However, the pholidosis of our lowland D. dryada completely falls within the variation of a large series of topotypical D. clarkorum from Cankurtaran pass (700 m). Con- sequently, we regard D. dryada to be a junior synonym of D. clarkorum.
Populations from warmer lowland localities tend to a finer pholidosis with relatively higher sca- le counts than populations from the cooler high elevation localities. This kind of variation re- minds of the principle of `numerical variation of pholidosis correlated with climate` described by SCHMIDTLER (1986) from similar data-sets in green lizard species of the genus Lacerta (sensu stricto). Therefore most of the variation is supposed to be modificatory and taxonomic con- sequences are rejected at present.
The rock lizard D. clarkorum occurs within the Euxinic zone with more than 1200 mm annual precipitation, from thermophilic coastal Colchian broad-leaved forest up to montane conifer forest. D. clarkorum may be associated with D. derjugini barani, D. parvula adjarica, D. rudis ssp., and Darevskia sp. The habitat spectrum of D. clarkorum seems to be narrow, if the spe- cies lives sympatrically with other rock-dwelling lizards. In such cases D. clarkorum is restric- ted to mostly shaded, moist-cool microhabitats. Without competition, however, the habitat spectrum becomes considerably broader, including even dry and rather exposed structures. Bischoff, W. (2003) - Die Eidechsenfauna Georgiens. Teil II. Die Gattung Darevskia. - Die Eidechse, Bonn, 14 (3): 65-93. × In Georgia, 16 species of rock lizards of the Genus Darevskia are occuring (D. alpina, D.
„armeniaca`, D. brauneri, D. caucasica, D. clarkorum, D. daghestanica, D. „dahli`, D.
derjugini, D. mixta, D. nairensis, D. parvula, D. portschinskii, D. praticola, D. rudis, D.
„unisexualis` and D. Valentini). Besides short presentations of the single species and hints on their distribution and habitats, also some systematic remarks are given. Ciobanu, D.G. & Grechko, V.V. & Darevsky, I.S. (2003) - Molecular Evolution of Satellite DNA CLsat in Lizards from the Genus Darevskia (Sauria: Lacertidae): Correlation with Species Diversity. - Russian Journal of Genetics, Moscow, 39 (11): 1292-1305. × The structure and evolution of a satellite DNA family was examined in lizards from the genus Darevskia(family Lacertidae). Comparison of tandem units of repeated DNA (satDNA), CLsat, in all species from the genus Darevskiahas shown that their variability is largely explained by single-nucleotide substitutions, which form about 50 diagnostic positions underlying classification of the family into three subfamilies. Maximum differences between the subfamilies reached 25%. At this level of tandem unit divergence in the subfamilies, no cross-hybridization between them was observed (at 65°C). The individual variability within one subfamily within the species was on average 5% while the variability between species consensuses within a subfamily was 10%. The presence of highly conserved regions in all monomers and some features of their organization show that satellites of all Darevskia species belong to one satDNA family. The organization of unit sequences of satellites CLsat and Agi160 also detected by us in another lizard genus, Lacerta s. str. was compared. Similarity that was found between these satellites suggests their relatedness and common origin. A possible pathway of evolution of these two satDNA families is proposed. The distribution and content of CLsat repeat subfamilies in all species of the genus was examined by Southern hybridization. Seven species had mainly CLsatI (83 to 96%); three species, approximately equal amounts of CLsatI and CLsatIII (the admixture of CLsatII was 2–5%); and five species, a combination of all three subfamilies in highly varying proportions. Based on these results as well as on zoogeographic views on the taxonomy and phylogeny of the Darevskia species, hypotheses on the evolution of molecular-genetic relationships within this genus are advanced. Чобану Д.Г., Гречко В.В., Даревский И.С. (2003) - Молекулярная эволюция сателлитной ДНК CLSAT ящериц рода Darevskia (Sauria: Laeertidae): корреляция с видовым разнообразием. - Генетика, 39 (11): 1527-1541. × Исследовали структуру и эволюцию семейства сателлитной ДНК ящериц рода Darevskia сем. Lac-ertidae. При сравнительном анализе последовательностей мономеров тандемных повторов ДНК (сатДНК) - CLsat - всех видов ящериц р. Darevskia показано, что вариабельность повторов обусловлена главным образом однонуклеотидными заменами, которые образуют около 50 диагностических позиций, на основании которых выделены три подсемейства повторов. Максимальные значения различий между подсемействами достигают 25%. При такой степени дивергенции мономеров трех подсемейств перекрестной гибридизации (при 65°С) между ними не наблюдается. Индивидуальная вариабельность мономеров одного подсемейства внутри вида в среднем составляет 5%, а вариабельность между видовыми консенсусами внутри подсемейства - 10%. Наличие высококонсервативных областей во всех мономерах и ряд особенностей их организации позволяют рассматривать сателлиты всех видов р. Darevskia как единое семейство сатДНК. Приведено сравнение организации последовательности мономеров сателлита CLsat и обнаруженного нами Agi160 из другого рода ящериц -Lacerta s. str. Выявлено сходство, свидетельствующее о родстве этих сателлитов и об их общем происхождении, предложен вероятный путь эволюции этих двух семейств сатДНК. Методом Саузерн-гибридизации изучено распространение и содержание подсемейств повторов CLsat во всех видах рода. Семь видов содержат в основном CLsatl (от 83 до 96%), три вида - примерно равные количества CLsatl и CLsatIII (примесь CLsatII составляет 2-5%), а пять видов - сумму всех трех подсемейств в сильно отличающихся соотношениях. На основе этих данных, в совокупности с зоогеографически-ми представлениями о родстве видов р. Darevskia, предложены гипотезы эволюции молекулярно-генетического родства видов этого рода. Ilgaz, C. (2007) - The Morphology, Taxonomy, and Distribution of Specimens of Darevskia clarkorum(Sauria: Lacertidae: Darevskia) Collected from Turkey’s Eastern Black Sea Region. - Turkish Journal of Zoology, Ankara, Turkey, 31 (4): 325-336. × In all, 103 specimens of Clark’s lizard, Darevskia clarkorum(Darevsky and Vedmederja, 1977), from 6 localities in the
eastern Black Sea region of Turkey were examined in terms of pholidosis characteristics, morphometric measurements, and color
and pattern features. Kolmogorov-Smirnov testing across all populations verified sexual dimorphism in the Ortacalar and Hemflin
populations, regarding transversal series of ventral plates (P < 0.05). According to the Mann-Whitney U test, 16 pholidosis
characteristics showed differences among the populations (P < 0.05). For comparison of morphometric measurements, 4
morphometric indices and ratios were developed. According to the results of independent t-testing, no differences were found
between males and females among the populations. One-way variance analyses based on 4 morphometric indices and ratios
confirmed that head index, snout-vent length/tail length, and tail length/total body length showed differences among the populations
(P < 0.05). Regarding pholidosis characteristics (except rates of the presence of a masseteric plate and contact between the rostral
and internasal plate), morphometric measurements, and color and pattern features, the specimens collected from 6 different
localities were similar to D. clarkorum specimens mentioned in the literature. Furthermore, the known range of the species in Turkey
has been extended.
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